By species www.diverosa.com Rokus Groeneveld & Sanne Reijs   
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Chromodoris or Goniobranchus?

In 2012 Rebecca Johnson and Terrence Gosliner published ‘Traditional taxonomic groupings mask evolutionary history: A molecular phylogeny and new classification of the chromodorid nudibranchs’. This had major consequences naming the chromodorid nudibranchs, and above all for the Chromodoris family, as this page proves. All these species were formerly known as Chromodoris species and have been reclassified as Goniobranchus.

For the complete publication of Rebecca Johnson & Terrence Gosliner click here.

FAN-367 Goniobranchus albonares, White gill goniobranchus

Goniobranchus albonares

White gill goniobranchus

Anilao, Philippines 2015
Dive site: Dive and trek

 FR2-116_Goniobranchus_albopunctatus

Goniobranchus albopunctatus

?

Romblon, the Philippines 2023
Dive site: Dive site: Bonbon end

 FRO-220_Goniobranchus_albopustulosus

Goniobranchus albopustulosus

?

Romblon, the Philippines 2017
Dive site: Logbon, Sanctuary

FR2-068_Goniobranchus_albopustulosus

Goniobranchus albopustulosus

?

Romblon, the Philippines 2023
Dive site: Dive site: Bangug check (night dive)

 EDS-29 Ringed goniobranchus, Goniobranchus annulatus

Goniobranchus annulatus

Ringed goniobranchus

Egypt, Deep South 2004
Dive site: Wadi gimal, Torfa

 IL3-051 Gold and Purple Goniobranchus, Goniobranchus aureopurpureusa

Goniobranchus aureopurpureus

Gold and Purple goniobranchus

Lembeh, Indonesia 2008
Dive site: Jarijari-Jahir (night dive)

 IBA-301 Gold and Purple goniobranchus, Goniobranchus aureopurpureus

Goniobranchus aureopurpureus

Gold and Purple goniobranchus

Bali, Indonesia 2009
Dive site: Seraya

 IB3-043_Goniobranchus_aureopurpureus,_Gold_and_purple_goniobranchus

Goniobranchus aureopurpureus

Gold and Purple goniobranchus

Bali, Indonesia 2016
Dive site: Puri jati

IL6-268_Goniobranchus_aureopurpureus

Goniobranchus aureopurpureus

Gold and Purple goniobranchus

Lembeh, Indonesia 2022
Dive site: Bianca

 ANS-345_Goniobranchus_aureopurpureus

Goniobranchus aureopurpureus

Gold and Purple goniobranchus

Australia, New South Wales 2023
Dive site: Nelson bay, Fly point

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Slugs & Flatworms 		EMA-072 Red sea or Charlott's goniobranchus, Goniobranchus charlottae

Goniobranchus charlottae

Red sea of Charlott's goniobranchus

Marsa alam, Egypt 2012
Dive site: house reef Oasis (night dive)

 MKA-26 Co goniobranchus, Goniobranchus coi

Goniobranchus coi

Co's goniobranchus

Kapalai, Malaysia 2003
Dive site: house reef (Mandarin valley)

IBA-011 Co goniobranchus, Goniobranchus coi

Goniobranchus coi

mantle flapping Co's goniobranchus

Bali, Indonesia 2009
Dive site: Menjangan, Pos I

 IBA-143 Co goniobranchus egg ribbon, Goniobranchus coi

Goniobranchus coi

Co's goniobranchus egg ribbon

Bali, Indonesia 2009
Dive site: Menjangan, Pos I

 IAM-157 Co goniobranchus, Goniobranchus coi

Goniobranchus coi

Co's goniobranchus

Ambon, Indonesia 2010
Dive site: Laha III

 IAM-140 Co goniobranchus, Goniobranchus coi

Goniobranchus coi

Co's goniobranchus

Ambon, Indonesia 2010
Dive site: Amahusa

 IHW-041 Co goniobranchus, Goniobranchus coi

Goniobranchus coi

Co's goniobranchus

Halmahera, Indonesia 2010
Dive site: N 00.28.192 / E 127.58.6668

IB2-137 Co goniobranchus, Goniobranchus coi

Goniobranchus coi

Co's goniobranchus

Bali, Indonesia 2011
Dive site: Amed, Japanese wreck

 FNE-016 Co goniobranchus, Goniobranchus coi, with egg ribbon

Goniobranchus coi

Co's goniobranchus laying eggs

Negros, Philippines 2012
Dive site: Siquijor, Paliton (cave)

 FM2-011 mantle flapping Co goniobranchus, Goniobranchus coi

Goniobranchus coi

mantle flapping Co's goniobranchus

Cebu, Philippines 2012
Dive site: Fish feeding

 IH2-037 Co goniobranchus, Goniobranchus coi

Goniobranchus coi

Co's goniobranchus

Halmahera, Indonesia 2014
Dive site: Tanjung uli

 FAN-016 Goniobranchus coi, Co goniobranchus

Goniobranchus coi

Co's goniobranchus

Anilao, Philippines 2015
Dive site: Red rock

Species of Goniobranchus can show 'mantle-flapping'. The whole mantle edge rises and falls simultaneously. For example Goniobranchus Coi and Gon. kuniei do that.
Gon. hintuanensis and Gon. geometrica show a different type of 'mantle-flapping'. Only the front of the mantle edge rises and falls in a rhythmic wave. There’s no idea what the 'purpose' is for mantle flapping, if there is any.
FRO-267_Goniobranchus_coi_Cos_goniobranchus

Goniobranchus coi

Co's goniobranchus

Romblon, the Philippines 2017
Dive site: Tobias corner

 IMB-192_Goniobranchus_coi

Goniobranchus coi

Co's goniobranchus

Manado Bay, Indonesia 2022
Dive site: Little beach

 IB4-010_Goniobranchus_coi

Goniobranchus coi

Co's goniobranchus

Bali, Indonesia 2024
Dive site: Bayu’s place

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Slugs & Flatworms
IBA-199 Collingwood's goniobranchus, Goniobranchus collingwoodi

Goniobranchus collingwoodi

Collingwood's goniobranchus

Bali, Indonesia 2009
Dive site: Tulamben, USAT Liberty

 IB4-073_Goniobranchus_collingwoodi

Goniobranchus collingwoodi

Collingwood's goniobranchus

Bali, Indonesia 2024
Dive site: Coral garden (night dive)

  ANS-050_Goniobranchus_daphne

Goniobranchus daphne

?

Australia, New South Wales 2023
Dive site: Clifton gardens (night dive)

 ANS-250_Goniobranchus_daphne

Goniobranchus daphne

?

Australia, New South Wales 2023
Dive site: Bare island

ANS-384_Goniobranchus_daphne

Goniobranchus daphne

?

Australia, New South Wales 2023
Dive site: Nelson bay, Pipeline

  ASA-057_Goniobranchus_epicurius

Goniobranchus epicurius

?

South Australia 2018
Dive site: Port Hughes jetty

  IH2-086 Reliable goniobranchus, Goniobranchus fidelis

Goniobranchus fidelis

Reliable goniobranchus

Halmahera, Indonesia 2014
Dive site: Mangrove (night dive)

IL5-045 Reliable goniobranchus, Goniobranchus fidelis

Goniobranchus fidelis

Reliable goniobranchus

Lembeh, Indonesia 2014
Dive site: Pantai parigi

 FAN-467 Goniobranchus fidelis, Reliable goniobranchus

Goniobranchus fidelis

Reliable goniobranchus

Anilao, Philippines 2015
Dive site: Basura

 IB3-309_Goniobranchus_fidelis,_Reliable_goniobranchus

Goniobranchus fidelis

Reliable goniobranchus

Bali, Indonesia 2016
Dive site: Tulamben, house reef Ocean view

 IB3-144_Goniobranchus_fidelis,_Reliable_goniobranchus

Goniobranchus fidelis

Reliable goniobranchus

Bali, Indonesia 2016
Menjangan, Mimpi chanel (night dive)

 FMO-024_Goniobranchus_fidelis

Goniobranchus fidelis

Reliable goniobranchus

Anda, Philippines 2018
Dive site: J. Edens place

  FMI-234 Geometric goniobranchus, Goniobranchus geometricus

Goniobranchus geometricus

Geometric goniobranchus

Cebu, Philippines 2007
Dive site: Kasai wall

 IGI-50 Geometric goniobranchus, Goniobranchus geometricus

Goniobranchus geometricus

Geometric goniobranchus

Gili Islands, Lombok, Indonesia 2009
Dive site: Meno wall, Gili Meno

 IL4-175 Geometric goniobranchus, Goniobranchus geometricus

Goniobranchus geometricus

Geometric goniobranchus

Lembeh, Indonesia 2010
Dive site: Batu lubang (night dive)

 IL4-204 Geometric goniobranchus, Goniobranchus geometricus

Goniobranchus geometricus

Geometric goniobranchus

Lembeh, Indonesia 2010
Dive site: Kapal indah

IB2-049 Geometric goniobranchus, Goniobranchus geometricus

Goniobranchus geometricus

Geometric goniobranchus

Bali, Indonesia 2011
Dive site: Kubu, Batu kembar

 FM2-204 Geometric goniobranchus, Goniobranchus geometricus

Goniobranchus geometricus

Geometric goniobranchus

Cebu, Philippines 2012
house reef Magic Island (night dive)

IMB-087_Goniobranchus_geometricus

Goniobranchus geometricus

Geometric goniobranchus

Manado Bay, Indonesia 2022
Dive site: MDJ

 ANS-128_Goniobranchus_geometricus

Goniobranchus geometricus

Geometric goniobranchus

Australia, New South Wales 2023
Dive site: Nelson bay, Fly point

 shortcut to pages:
Chromodoris
Hypselodoris
other Chromodorididae
Discodorididae
Facelinidae
Phyllidiidae
Polyceridae
other nudibranches
IL3-055 Lumpy goniobranchus, Goniobranchus hintuanensis

Goniobranchus hintuanensis

Lumpy goniobranchus

Lembeh, Indonesia 2008
Dive site: Police pier II

 IBA-245 Lumpy goniobranchus, Goniobranchus hintuanensis

Goniobranchus hintuanensis

Lumpy goniobranchus

Bali, Indonesia 2009
Dive site: Tulamben, Drop off

 IB2-257 Lumpy goniobranchus, Goniobranchus hintuanensis

Goniobranchus hintuanensis

Lumpy goniobranchus

Bali, Indonesia 2011
Dive site: Tulamben, USAT Liberty

 IB2-110 Lumpy goniobranchus, Goniobranchus hintuanensis

Goniobranchus hintuanensis

Lumpy goniobranchus

Bali, Indonesia 2011
Dive site: Seraya

 FAN-163 Goniobranchus hintuanensis, Lumpy goniobranchus

Goniobranchus hintuanensis

Lumpy goniobranchus

Anilao, Philippines 2015
Dive site: Secret garden

FAN-469 Goniobranchus hintuanensis, Lumpy goniobranchus

Goniobranchus hintuanensis

Lumpy goniobranchus

Anilao, Philippines 2015
Dive site: Pinnacle

 FRO-193_Goniobranchus_hintuanensis_Lumpy_goniobranchus

Goniobranchus hintuanensis

Lumpy goniobranchus

Romblon, the Philippines 2017
Dive site: Bangug 3 (night dive)

 IMB-097_Goniobranchus_hintuanensis

Goniobranchus hintuanensis

Lumpy goniobranchus

Manado Bay, Indonesia 2022
Dive site: MDJ

 IL6-556_Goniobranchus_hintuanensis

Goniobranchus hintuanensis

Lumpy goniobranchus

Lembeh, Indonesia 2022
Dive site: Pulau makawide

 
IL3-079 Kuni’s nudibranch, Goniobranchus kuniei

Goniobranchus kuniei

Kuni’s nudibranch

Lembeh, Indonesia 2008
Dive site: Nudi retreat

 IHW-008 Kuni’s nudibranch, Goniobranchus kuniei

Goniobranchus kuniei

Kune's nudibranch

Halmahera, Indonesia 2010
Dive site: Shark reef

 IHW-132 Kuni’s nudibranch, Goniobranchus kuniei

Goniobranchus kuniei

Kuni’s nudibranch

Halmahera, Indonesia 2010
Dive site: Shark reef

 IHW-259 Kuni’s nudibranch, Goniobranchus kuniei

Goniobranchus kuniei

Kuni’s nudibranch

Halmahera, Indonesia 2010
Dive site: Gemaf

 IH2-031 Kuni’s nudibranch, Goniobranchus kuniei

Goniobranchus kuniei

Kuni’s nudibranch

Halmahera, Indonesia 2014
Dive site: Pasir putih (night dive)

IRA-061 Kunis nudibranch, Goniobranchus kuniei

Goniobranchus kuniei

Kuni’s nudibranch

Raja ampat, Indonesia 2014
Dive site: Yenbuba jetty

 FAN-172 Goniobranchus kuniei, Kuni's nudibranch

Goniobranchus kuniei

Kuni’s nudibranch

Anilao, Philippines 2015
Dive site: Matu (night dive)

 IB3-107_Goniobranchus_kuniei,_Kuni's_nudibranch

Goniobranchus kuniei

Kuni’s nudibranch

Bali, Indonesia 2016
Tulamben, between Drop off / Coral garden

 IMB-047_Goniobranchus_kuniei

Goniobranchus kuniei

Kuni’s nudibranch

Manado bay, Indonesia 2022
Dive site: Celebes

 
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Slugs & Flatworms 		IHW-083 Leopard chromodoris, Chromodoris leopardus

Goniobranchus leopardus

Leopard goniobranchus

Halmahera, Indonesia 2010
Dive site: 'Kangelang' jetty

 IH2-145 Leopard chromodoris, Chromodoris leopardus

Goniobranchus leopardus

Leopard goniobranchus

Halmahera, Indonesia 2014
Dive site: Gemaf muck

 IL5-184 Leopard chromodoris, Chromodoris leopardus and Elegant flatworm, Pseudobiceros bedfordi

Goniobranchus leopardus

Leopard goniobranchus

Lembeh, Indonesia 2014
Dive site: Horse shoe (night dive)

 IAM-348 Precious goniobranchus, Goniobranchus preciosus

Goniobranchus preciosus

Precious goniobranchus

Ambon, Indonesia 2010
Dive site: Devion point

IB3-218_Goniobranchus_preciosus,_Precious_goniobranchus

Goniobranchus preciosus

Precious goniobranchus

Bali, Indonesia 2016
Dive site: Secret bay (night dive)

  IL3-233 Red reticulate nudibranch, Goniobranchus reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Lembeh, Indonesia 2008
Dive site: Police pier

 IAM-327 Red reticulate nudibranch, Goniobranchus reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Ambon, Indonesia 2010
Dive site: Jetty

 FNE-113 Red reticulate nudibranch, Goniobranchus reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Negros, Philippines 2012
Dive site: Paliton sanctuary

FNE-136 Red reticulate nudibranch, Goniobranchus reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Negros, Philippines 2012
Dive site: Bahura

 IL5-286 Reticulated chromodoris, Goniobranchus reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Lembeh, Indonesia 2008
Dive site: Pintu kolaba

 IRA-237 Reticulated chromodoris, Goniobranchus reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Raja ampat, Indonesia 2014
Dive site: Kri south

 IB3-174_Goniobranchus_reticulatus,_Red_reticulate_nudibranch

Goniobranchus reticulatus

Red reticulate nudibranch

Bali, Indonesia 2016
Menjangan, Mimpi chanel (night dive)

 FRO-055_Goniobranchus_reticulatus_Red_reticulate_nudibranch

Goniobranchus reticulatus

Red reticulate nudibranch

Romblon, the Philippines 2017
Dive site: Tobias corner

FN2-197_Goniobranchus_reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Negros, Philippines 2019
Dive site: Masaplod norte

 IMB-305_Goniobranchus_reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Manado Bay, Indonesia 2022
Dive site: Celebes

 IL6-401_Goniobranchus_reticulatus

Goniobranchus reticulatus

Red reticulate nudibranch

Lembeh, Indonesia 2022
Dive site: Tanjung kubur

  ANS-153_Goniobranchus_roboi

Goniobranchus roboi

?

Australia, New South Wales 2023
Dive site: Bare island

FAN-186 Goniobranchus rubrocornutus

Goniobranchus rubrocornutus

?

Anilao, Philippines 2015
Dive site: Secret garden

  ANS-038_Goniobranchus_splendidus

Goniobranchus splendidus

?

Australia, New South Wales 2023
Dive site: Bare island

 ANS-079_Goniobranchus_splendidus

Goniobranchus splendidus

?

Australia, New South Wales 2023
Dive site: Nelson bay, Fly point

 AVI-009_Goniobranchus_tinctorius_Red-netted_goniobranchus_1

Goniobranchus tinctorius

Red-netted goniobranchus

Australia, Victoria 2018
Dive site: Blairgowrie pier

AVI-192_egg_ribbon_Goniobranchus_tinctorius

Goniobranchus tinctorius

egg ribbon Red-netted goniobranchus

Australia, Victoria 2018
Dive site: Blairgowrie pier

 AWA-204_Goniobranchus_tinctorius_1

Goniobranchus tinctorius

Red-netted goniobranchus

Western Australia 2020
Dive site: Ammo jetty

 AWA-218_Goniobranchus_tinctorius_2

Goniobranchus tinctorius

Red-netted goniobranchus

Western Australia 2020
Dive site: Ammo jetty (night dive)

 AWA-508_Goniobranchus_tinctorius

Goniobranchus tinctorius

Red-netted goniobranchus

Western Australia 2020
Dive site: Kwinana grain terminal

 EDS-28 Red-netted goniobranchus, Goniobranchus tinctorius

Goniobranchus tinctorius?

Red-netted goniobranchus?

Egypt, Deep South 2004
Dive site: Wadi gimal, Torfa

AWA-170_probably_Goniobranchus_tinctorius

Goniobranchus tinctorius?

Red-netted goniobranchus

Western Australia 2020
Dive site: Ammo jetty (night dive)

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Slugs & Flatworms 		IBA-246 Verrier's Goniobranchus, Goniobranchus verrieri

Goniobranchus verrieri

Verrier's goniobranchus

Bali, Indonesia 2009
Dive site: Tulamben, Drop off

 FN2-180_Goniobranchus_verrieri

Goniobranchus verrieri

Verrier's goniobranchus

Negros, Philippines 2019
Dive site: Pyramids

 AWA-148_Goniobranchus_sp

Goniobranchus sp.

?

Western Australia 2020
Dive site: Coogee marina, north wall